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本书以基本的符号意义机制为起点，阐述了符号的指示性，并对文化的"标出性"概念做了系统的分析讨论，进而从跨学科的角度探寻符号与生物符号的互动关系，，揭示了二者的符号意义，将分属自然与文化的两个议题在符号学的视阈中得以统一。
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