How did ancestral arthropods successfully diversify to colonize the land? How do insects use their wings to exploit terrestrial and freshwater ecosystems by metamorphosis? The causes and consequences of metamorphosis are still poorly understood in biology. This collection of essays examines the molecular basis underlying the metamorphosis regulation of the white wax scale insect (WWS), a fascinating insect that undergoes different metamorphoses between males and females. It shows that the WWS has produced great economic or medical benefits for human beings for thousands of years. This book provides interesting information for scientific and technical personnel, as well as undergraduate and graduate students engaged in entomology, molecular biology and biogeography.