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運命に導かれるままに、草食ライオンのレオンと仲間たちは紆余曲折ありながらも、ついに動物の都ライオネスクに辿り着いた。その都では宮廷や王族、人間たちの陰謀が複雑に絡み合っていた......。レオンの行く先にはさらなる大きな壁が立ちはだかる！
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