Madison Riley and Art Thompson are both members of a super secret group of street racers, American men and women who love the thrill of a cross country race and the chance to win a race pot every year worth millions of dollars. They are both members of the Fireball for Life Club, having placed in highly illegal races in the last ten years. Fireballers, as they are called, drive red sports cars for the events.

This year's Fireball race goes from Miami to Seattle. The race sponsor decided that the drivers would be in vintage convertibles for the trip, meaning that Art would need to dust off his 1966 Shelby Cobra for the trip. Partnerless this year, as his usual partner is laid up with an injury, Art chooses Madison, or Maddie, as his partner for the 3,300 mile race as her usual partner is conveniently on maternity leave and she is the only one of the partnerless drivers he trusts not to wreck his Cobra.

Normally, Art and Maddie avoid each other like the plague both during race week, and on the social circuit in Manhattan, but forced together for lead up to the race, they discover more than they ever thought possible about each other. Along the way, the colorful cast of characters among the racers add to the drama of race week as old relationships dissolve and new ones take their place.