A lone survivor in a zombie-infested world, Jonah Caine wandered for months, struggling to understand the apocalypse in which he lives.
Unable to find a moral or sane reason for the horror that surrounds him, he is overwhelmed by violence and insignificance. Then Jonah comes across a group of survivors living in a museum-turned-compound. They are led by Jack, an ever-practical and efficient military man; and Milton, a mysterious prophet who holds a strange power over the dead. Both share Jonah's anguish over the brutality of their world as well as his hope for its beauty. Together with others, they build a community that reestablishes an island of order and humanity surrounded by relentless ghouls. But this newfound peace is short-lived, as Jonah and his band of refugees clash with another group of survivors who remind them that the undead are not the only—nor the most grotesque—horrors they must face.