Lioness Cristabel Hardison will do anything to avoid returning to Kentucky, where her father, the king of their pride, wants to mate her to a male of his choosing. When her roommate has to vacate their condo, Cris seizes the chance to stay with her best friend, Lily, in Ashland, hoping to find a new job and a place to live.

Cousins Dylan, Chase, and Hunter Hall have given up hope of ever finding a mate to share. While other lions have found their mates, the cousins believe they might be destined to remain single. But one day, at the town diner, they catch the scent of their mate, and are determined to find her no matter what.

When Cris meets her mates, sparks fly instantly, and she becomes the center of their world. However, her father isn't pleased. When he brings an uninvited guest to crash her newfound happiness, Cris realizes that her future—and her mates'—may be in jeopardy.

Will Cris and her mates get the happily ever after they all long for, or will her father's schemes tear them apart?