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一年一度的南遷越冬即將開始，天鵝隊長伯納德正加緊訓練他的團隊。他們的目的地是3,000公里以外的美國亞利桑那州。遷徙途中，危險重重──上有禿鷲，下有狐狸，而且路途遙遠，實不容有失。這時候，偶然救了他們的小松鼠丹尼向伯納德提出一個建議。只是這一改變，令天鵝遇到更大的危險......他們能化險為夷嗎？
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