Life on earth has changed forever, as humans and their courageous robotic allies, the Autobots, must warily work together to protect the planet from the destructive forces of the evil Decepticons. At the headquarters of NEST, tech sergeant Epps and captain Lennox both guard and assist cyberneticist Kaminari Ishihara and the brooding Russian AI genius Petr Andronov as they explore the differences between organics and bots. All around them, alliances fray, distrust grows, suspicions mount, and traitors come out of the shadows.

Meanwhile, Optimus Prime, the powerful leader of the Autobots who is also part of NEST, plays defense, as battles flare up from Australia to Zambia. But escalating Decepticon attacks will culminate in a final confrontation from which no one--man or machine--will emerge unscathed.