When Annabelle loses her first tooth, she sparks a movement in Toothland, a magical mouse kingdom that relies on clean teeth for power. The kingdom thrives under King Molar, Lady Flossina, Sir Canine, and Princess Pearl as children's brushing habits fuel its energy. However, Snaggles, feeling excluded, unleashes a Sugar Storm that threatens Toothland. As chaos spreads and Toothland falters, its defenders—including Annabelle—work together to restore good dental habits among children. In the final confrontation, Annabelle convinces Snaggles to change his ways, turning him into an advocate for dental care. Toothland is restored, united by healthy habits, and continues to shine thanks to the efforts of children everywhere.