Bruce's grandpa, the oldest butterfly and village patriarch, leaves home to find the fabled Butterfly Paradise. Bruce's father escorts him, suspecting this "paradise" might be something else altogether. Meanwhile, in the village, dissension arises over the new methods for harvesting and sharing nectar. When Bruce's uncle hires wasps to control the unhappy butterflies, Bruce and his mother endorse rebellion. Without Dad and Grandpa, though, they cannot prevail. Bruce gathers his friends to help and hurries to Butterfly Paradise. There, he is stunned to discover his relatives are prisoners. Worse, Grandpa is sick, and Bruce must find a way to rescue his family before it is too late.